Volunteer Research Projects on the Cedar Mountain Formation

by Angela Matthias

 

A NEW BASAL HADROSAUR FROM THE LOWER CRETACEOUS

CEDAR MOUNTAIN FORMATION OF EASTERN UTAH

David Gilpin and Tony DiCroce

            A new hadrosaur from the Lower Cretaceous Yellow Cat Member of the Cedar Mountain Formation was discovered in May of 2001.  This taxon is represented by a well-preserved left ilium, preacetabular process of the right ilium, right tibia, right metatarsal III, fused sacrum, ribs, and ossified tendons.  The ilium overall resembles that of the iguanodontid Camptosaurus, but it has the lateral process (“antitrochanter”) that characterizes hadrosaurs.  Plesiomorphic character of the elements suggests a recent departure from iguanodontids.  We conclude therefore that this taxon represents a basal

hadrosaur. 

 

A SAUROPOD FROM THE LOWER RUBY RANCH MEMBER OF THE CEDAR MOUNTAIN FORMATION OF EASTERN UTAH

David Gilpin, Peggy LeMone and John Scandizzo

               Sauropods from the Ruby Ranch Member of the Cedar Mountain Formation have only been known from the upper layers of this Member.  In May of 1999 the remains of a sauropod from the lower portion of the Member was discovered.  The bones of the animal were removed from the ground over a period of three visits to the site.  Their identity was unknown at that time.  Preparation, including assembly of surface scattered fragment s, however, has since revealed the specimen to be pelvic bones (two pubes and one ischium) of a sauropod Other bones consist of parts of long bones, ribs and other unidentifiable pieces.  The ends of the pubes are missing, making it unlikely this dinosaur can be identified more specifically. The stratigraphic position of this animal, low in the Ruby Ranch and the other sauropod specimens high in the Member suggests that they were continuous members of the Utah dinosaur fauna throughout this time frame. 

 

A LARGE NODOSAURID ANKYLOSAUR FROM THE

CEDAR MOUNTAIN FORMATION OF UTAH

David  Warren

Nodosaurs are well known from the Cretaceous of North America, with Sauropelta the best known Early Cretaceous form from the Cloverly Formation in Wyoming and Montana. Fragmentary remains of a huge nodosaurid better matched to Saropelta than any other ankylosaur were found weathered out of the Ruby Ranch Member of the Cedar Mountain Formation in eastern Utah.  The most describable elements are a very large right humerus and two very large cervical spikes; other identifiable elements include a fibula, centra of dorsal vertebra, and two small scutes. The ~65 cm long humerus indicates an individual approximately 6.8 m long, making it one of the largest known nodosaurids. The presence of Sauropelta supports previous interpretations that the upper portions of the Ruby Ranch Member are equivalent to parts of the Cloverly based on the presence of the iguanodontid Tenontosaurus. Such a correlation implies that the Ruby Ranch Member is in part Aptian-Albian in age.

 

GASTROLITHS FROM THE LOWER CRETACEOUS SAUROPOD

CEDAROSAURUS WEISKOPFAE

Frank Sanders (and Kim Manley)

A set of 115 clasts, ranging in size and mass from 0.04 cc to 270 cc and 0.1 gm to 715 gm, has been collected in association with a newly described brachiosaurid in the Cedar Mountain Formation, Cedarosaurus weiskopfi. The clasts were partially matrix-supported, and some were supported on-edge. There were a number of clast-to-clast and clast-to-bone contacts. The clast are most parsimoniously interpreted as gastroliths, making this the first set discovered in-situ in this formation. The gastrolith surfaces are mostly polished. Tight spatial distribution, partial matrix support, and some instances of on-edge orientation indicate that they were deposited while contained within carcass soft tissue. Low-energy depositional conditions and apparent initial containment within soft tissue indicate that the set is complete. Surface characteristics and distributions of shape, mass, volume, and composition have been determined. More than half of the clasts are less than 10 cc in volume. Drab colors indicate low selectivity by the sauropod for this characteristic. Most of the gastroliths are chert or quartzite, but some are sandstones and siltstones. High surface reflectance values of the majority of the gastroliths is consistent with the results of previous studies on other bona fide gastroliths.

 

Articulated Dermal Armor of  the ankylosaur Gastonia

Kathleen Brill

A large slab of articulated dermal armor of Gastonia, a polacanthine dinosaur, was found at Lorrie’s Site, a monospecific bonebed in the Cedar Mountain Formation (Lower Cretaceous) in east-central Utah. This find is significant because armor pattern is poorly known in ankylosaurs. The osteoderms are arranged in a distinctive series of interconnected rosettes consisting of large osteoderms surrounded by rings of smaller elements, closely fitted together like tiles.  The largest osteoderms (35 to 55 mm diameter) are roughly elliptical with a short peak that are typically placed off-center. The peaks are asymmetrical and pitted.  The large osteoderms often have faint grooves, which radiate from the peaks to the margins. The smaller osteoderms (approximately 6 to 25 mm) are irregularly shaped polygons with straight margins where they interface with a neighboring element.  The smaller osteoderms have flat, parallel top and base surfaces with a rough texture.

            The armor sheet was preserved with the external surface up.  It was found in mudstone, which directly overlaid a hard concretion layer, approximately 20 to 30 cm thick, which encased the bones of Gastonia. The sheet is undulating, conceivably indicating that the skin was flexible when deposited.  Cervical ribs, dorsal ribs, vertebra fragments, ossified tendons, and a single ulna are adjacent to the armor.  This, and the absence of larger armor elements, could signify that this skin was from the anterior of the animal. 

 

A poster describing the work thus far was presented at the 2004 Society of Vertebrate Paleonotology meeting in Denver Colorado.  Preparation of the bone found in the concretions beneath the armor slab is currently in-process with the goal of finding more armor.  Future work will concentrate on identifying additional specimens of articulated armor, discerning the armor patterns, and the describing the locations of  armor within  the skin of the animal.

 

CORTICATED PRESSURE EROSIONS, “PITTING”,

IN OSTEODERMAL ANKYLOSAUR ARMOR

Lorrie A. McWhinney and Angela Matthias

The osteodermal armor of ankylosaurs often exhibits a pattern of depressions that are clearly not associated with taphonomic alteration.  An investigation of this pathology was initiated using Gastonia osteodermal armor material primarily from the Lorrie’s Bone Bed Site in the Ruby Ranch Member of the Early Cretaceous Cedar Mountain Formation, eastern Utah. The site is a monospecific bonebed of the polacanthid Gastonia with a minimum of a dozen individuals, possibly more.  Additional osteodermal armor material from other collections was also used for correlation.  The internal wall of each depression is lined with cortical bone indicating an ante-mortem osteoblastic response to an unknown disease process.  This one characteristic is useful in distinguishing this pathology from a non-pathologic, post-mortem alteration as seen in exposed, eroded trabecular bone. Based on cortical erosion in modern bone (human and animal), probable differential etiologies include; foreign body granuloma, intraosseous epidermoid inclusion cyst, fungal and /or bacterial infection, cutaneous dyskeratosis and glomus tumor.  Independent of the etiology, the “pits” on the osteodermal material should be referred to as “corticated pressure erosions”.

 

A New Species of Gastonia (Dinosauria: Ankylosauria)

from the Cedar Mountain Formation of Eastern Utah

BILLY KINNEER

A mass graveyard of several disarticulated skeletons of a new species of the ankylosaur Gastonia has been discovered near the base of the Ruby Ranch Member of the Cedar Mountain Formation. The site designated as Lorrie’s Site is located approximately 13 km north of the Arches National Park and 52 km east of Green River in Grand County, Utah.

            To date over 3000 bones representing at least 20 individual animals have been collected and prepared from this monospecific ankylosaur bonebed.  Over 75 skull fragments representing at least 9 individuals are available for study.  Numerous fragments of scapula and armor, as well as other complete and fragmented post cranial bones, are available for comparison.  The differences discovered in the examination of the features associated with the skull, scapula and amour strongly suggests that this is a new species of Gastonia.

            The importance of this discovery further extends the support for ankylosaurs being present in North America during the Early Cretaceous, but more importantly it now proves there was more ankylosaur diversity than was previously understood.

 

RECONSTRUCTION OF THE PELVIC AND HIND LIMB

MUSCULATURE IN THE ANKYLOSAUR GASTONIA

Tony DICROCE and Billy KINNEER

There have been many previous attempts to describe the ankylosaur pelvic musculature structure as a basis for explaining and understanding the evolution and development of its unusual ilium. All previous attempts have been incomplete; either important muscles were not fully considered or the skeletal elements were incomplete. None of the previous studies offered a convincing argument to explain the functionality of the ilium.  Unlike most other attempts at reconstruction, this model shows the superficial and deep muscles in full-size three-dimension. A cast of an ankylosaur pelvic skeletal girdle was constructed from material recovered from an extensive burial of several individual animals, possibly a new species of Gastonia, in the Cedar Mountain Formation in eastern Utah. The muscles were identified based on dissection of birds (Gallus, Megeagris) and crocodile (Caiman), sculpted in clay on the pelvic skeletal. Muscle scars on the original elements were interpreted for origin and insertion points. The broad flat ilium blade is the origin for the large tricep extensors, M. ilio-tibialis, along the lateral margin. The ischium has developed proximally and with the synsacrum forms an anteromedial wall along the ilium medial margin making up the enclosed shallow acetabulum. The pubis has a thin, blade shaped, postpubic process and retains muscle scars suggesting platforms for M. ambiens and M. pubo-ischio femoralis. The location of M. ambiens in this reconstruction is placed anterolaterally to an extreme preacetabular position, unlike any other ornithischian.

            Additional studies with this model will be made to determine, what if any affect, dermal armor, locomotion, or diet had on the development of the anklyosaur pelvic structure. We anticipate this model will provide insight to the development and functionality of the ankylosaur ilium.

 

Evolutionary relationships of Early Cretaceous

Sauropod Faunas from the Cedar Mountain

Formation of Utah

Virginia Tidwell

Although sauropod dinosaurs are well known from the Upper Jurassic Morrison Formation sediments found  throughout the Western interior of the North America, until recently little was known about more recent, Cretaceous sauropods in the U.S.  However, several sauropod skeletons have recently been excavated  by DMNH crews from the Lower Cretaceous Cedar Mountain Formation (CMF) of Utah. These skeletons come from several different stratigraphic levels within the formation, and appear to represent differing taxa.  Identifying each species is the first step towards understanding the overall faunal diversity within the Cedar Mountain Formation. This diversity can then be correlated with Cretaceous sauropod assemblages worldwide.

            The bones from these DMNS sauropod quarries in the CMF were brought to the museum for cleaning and repair.  I have compared them with bones from the well known Late Jurassic sauropods Camarasaurus, Diplodocus and Brachiosaurus in order to determine to which group the new specimens are most closely related.  Additional comparisons were made with another sauropod group known as titanosaurs, that, although poorly represented in North American sediments, are well known from other parts of the world.  These comparisons required visiting museums housing major dinosaur collections in the U.S.(including AMNH, CM, USNM, and the Field Museum), Argentina (six museums), and England (two museums). 

            Partial skeletons from eight CMF localities have been identified to date. The results of these efforts so far suggest that one of these new CMF sauropods, Cedarosaurus weiskopfae (Tidwell et al 1999) resembles  Brachiosaurus more than any other known sauropod, although differing in several important areas.  Another new specimen, Venenosaurus dicrocei (Tidwell et al 2001), is a bit more like a titanosaur.  One partial skeleton composed of several posterior cervical vertebrae from the Yellow Cat member of the CMF has tentatively been  identified as a titanosaur due to the very low neural spines on all the elements, a titanosaur characteristic.  Three partial skeletons from the Cedar Mountains Ruby Ranch member also appear to represent titanosaurs.  The unexpected recovery of a sauropod forelimb from the base of the Dakota Formation suggests that titanosauriform sauropods were still present in the early Late Cretaceous, and represent the first verifiable sauropod specimen from the Dakota of Utah.  Additional material from the Poison Strip and Yellow Cat member are still under study.

 

Worldwide significance of Early Cretaceous

Titanosauriform dinosaurs

Virginia Tidwell

            In recent years new discoveries of titanosauriformes in North America, Europe and South America have fueled a significant expansion in Early Cretaceous sauropod research. Our study of previously described taxa, as well as many new specimens, seeks to provide detailed correlations with sauropods throughout these regions, and also develop rigorous time constraints for the North American titanosauriform radiation.  One of our current projects focuses on a large skeletal assemblage of Early Cretaceous sauropods that was excavated by the University of Utah in the 1970's at the Long Walk Quarry in the Cedar Mountain Formation of Utah.

            The Denver Museum is now working in cooperation with the U. of Utah to prepare a great many bones in over 20 field jackets that remain to be prepared, work that is slowed by the extremely hard matrix in which these bones are lodged.  We expect to continue the taxonomic study of these specimens, utilizing information gained by one of us (VT) in previous visits to many museums in the U.S., Argentina and England.  Recently we have begun to correlate these and other Early Cretaceous sauropod discoveries from Utah with similar finds from Argentina and England (Carpenter et al 2002; Tidwell and Carpenter 2002). We also hope to expand these correlations to other global areas with an in-depth study of the holotypic elements of  the titanosaur Ampelosaurus in Esparaza, France, and of the African Brachiosaurus housed in the Museum fur Naturkunde der Humbolt-Universitat, Berlin.

            Premilinary results suggest that the Long Walk specimen is more closely related to titanosaurs than to Brachiosaurus. This means that it is not the North American brachiosaur Cedarosaurus, nor Pelorosaurus, which is known from Wealden sediments in southern England.   The Long Walk specimen appears to more closely resemble basal titanosaurs such as Andesaurus or Epachthosaurus from Argentina.  It does not appear to be as derived as the African Malawisaurus, or similar titanosaurs from Argentina, although much preparation, and further on-site studies of holotypic specimens will be necessary before a final determination can be made.